Ayu Savitri Nurinsiyah
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The malacofauna of Java has been most studied among the Indonesian islands, but the list of land snails in the area remains outdated. This study presents an updated check list of land snails in Java and its adjacent islands. This list is a compilation data from field work in Java conducted in 2013-2016, records from various museums in Europe and Indonesia, collections from private collectors, data from citizen sciences, and literatures. In total, 263 land snail species were recorded in Java and its adjacent islands. The number comprises of 36 families i.e. Subclass Neritimorpha (2 families), Caenogastropoda (6 families), and Heterobranchia (28 families). About 40% are species endemic to Java and among them have restricted distribution to certain areas. In addition, 5% or 13 introduced species were recorded in Java.


biodiversity, Gastropoda, Indonesia, Mollusca, terrestrial

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Badan Pusat Statistik (BPS). 2017.

Bouchet, P., Rocroi, J.P., Hausdorf, B., Kaim, A., Kano, Y., Nützel, A., Parkhaev, P., Schrödl, M. & Strong, E.E. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia, 61(1–2): 1–526.

Butot, L. 1955. The molluscs fauna of Pulau Panaitan (Prinseneiland). Land and freshwater molluscs. Treubia, 23: 70–135.

Cannon, C.H., Morley, R.J. & Bush, A.B.G. 2009. The current refugial rainforests of Sundaland are unrepresentative of their biogeographic past and highly vulnerable to disturbance. PNAS, 106 (27): 11188–11193.

Dharma, B. 1992. Siput dan Kerang Indonesia II (Indonesian Shells II). Wiesbaden: Verlag Christa Hemmen [Indonesian].

Dharma, B. 2005. Recent & Fossil Indonesian Shells. Hackenheim: ConchBooks.

Dharma, B. 2007. Report on fossil Amphidromus and description of new species and new subspecies of recent and fossil Amphidromus from Indonesia (Gastropoda, Pulmonata: Camaenidae). Schriften zur Malakozoologie, 23: 45–78.

Dharma, B. 2014. A new species of Landouria Godwin-Austen 1918 from Jawa, Indonesia (Gastropoda, Bradybaenidae). Schriften zur Malakozoologie, 28: 7–12.

Dharma, B. 2021. New species and subspecies of recent and fossil Amphidromus Albers, 1850 from Java and Sumatra (Gastropoda: Camaenidae). The Festivus, 53(4): 298–306

Douglas, D.D., Brown, D.R. & Pederson, N. 2013. Land snail diversity can reflect degrees of anthropogenic disturbance. Ecosphere, 4 (2): 1–14.

Gomes, S.R. & Thomé, J.W. 2004. Diversity and distribution of the Veronicellidae (Gastropoda: Soleolifera) in the Oriental and Australian biogeographical regions. Memoirs of the Queensland Museum, 49: 589–601.

Greķe, K. 2019. New species and record of Diplommatina Benson, 1849 (Gastropoda: Diplommatinidae) from Java, Indonesia. Nautilus, 133: 14–21.

Hausdorf, B. 2019. Beyond Wallace’s line – dispersal of Oriental and Australo-Papuan Land-Snails across the Indo-Australian Archipelago. Zoological Journal of the Linnean Society, 185: 66–76.

Heryanto. 2008. Ekologi keong darat di Taman Nasional Gunung Ciremai. Jurnal Biologi Indonesia, 4: 359–370.

Heryanto. 2011. Land Snails of Java. Bogor: LIPI Press [Indonesian].

Heryanto, 2012. Keanekaragaman keong darat di dua macam habitat makro di Gunung Slamet Jawa Tengah. In: Maryanto, I., Noerdjito, M. & Partomihardjo, T., eds. Ekologi Gunung Slamet: Geologi, Klimatologi, Biodiversitas dan Dinamika Sosial. Jakarta: LIPI Press [Indonesian].

Heryanto. 2017. The Land snail diversity in succession forest of Mount Galunggung and stable forest of Mount Sawal, West Java. Zoo Indonesia, 26(2): 59–69.

Heryanto, Ristiyanti, M.M., Munandar, A. & Susilowati, P. 2003. Keong dari Taman Nasional Gunung Halimun, Sebuah Buku Panduan Lapangan. Cibinong: Biodiversity Conservation Project-LIPI-JICA-PHKA [Indonesian].

Hughes, A. 2017. Understanding the drivers of Southeast Asian biodiversity loss. Ecosphere, 8(1): 1–33.

Loosjes, F.E., 1953. Monograph of the Indo-Australian Clausiliidae (Gastropoda, Pulmonata, Clausiliidae, Phaedusinae). Beaufortia, 31: 1–226.

Martens, E. von, 1867. Die landschnecken. In: Die Preussische expedition nach Ost-Asien. Nach amtlichen Quellen. Zoologischer Theil. Vol. 2. Berlin: Decker [German].

Marwoto, R.M. 2011. Keong darat dan air tawar dari pulau Nusa Kambangan (Moluska, Gastropoda) (The terrestrial and freshwater snails from Nusa Kambangan Island (Mollusk, Gastropod)). Jurnal Moluska Indonesia, 2(1): 39–47.

Ministry of Environment and Forestry (KHLK). 2016. Statistik Kementrian Lingkungan Hidup dan Kehutanan Tahun. 2015. Jakarta: Pusat Data dan Informasi Kementrian Lingkungan Hidup dan Kehutanan Republik Indonesia [Indonesian].

Möllendorff, O. F. von. 1897. Neue Landschnecken von Java. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 29 (5/6):57–72 & (7/8): 89–97.

Mujiono, N. & Priawandiputra, W. 2020. Land snail from limestone karst in Java: an update with motes on their distribution. IOP Conf. Series: Earth and Environmental Science, 457(012002): 1–9.

Nurhayati, P.A., Affandi, M. & Nurinsiyah, A.S. 2021. Diversity and abundance of terrestrial gastropods on the slopes of Mount Arjuna-Welirang, East Java, Indonesia. Biodiversitas, 22(10): 4193–4202.

Nurinsiyah, A.S. 2015. Land snail fauna of the Sukolilo karst in Java (Indonesia). American Conchologist, 43: 30–32.

Nurinsiyah, A.S., Fauzia, H., Hennig, C. & Hausdorf, B. 2016. Native and introduced land snail species as ecological indicators in different land use types in Java. Ecological Indicators, 70: 557–565.

Nurinsiyah, A.S. & Hausdorf, B. 2017a. Revision of the Diplommatinidae (Gastropoda: Cyclophoroidea) from Java. Zootaxa, 4312: 201–245.

Nurinsiyah, A.S. & Hausdorf, B. 2017b. Dicharax (?) candrakirana n. sp. (Gastropoda: Cyclophoridae) from Sempu Island, Indonesia. Zootaxa, 4363(4): 589–591.

Nurinsiyah, A.S. & Hausdorf, B. 2019. Listing, impact assessment and prioritization of introduced land snail and slug species in Indonesia. Journal of Molluscan Studies, 85: 92–102.

Nurinsiyah, A.S., Neiber, M.T. & Hausdorf, B. 2019. Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java. European Journal of Taxonomy, 526: 1–73.

Nurinsiyah, A.S. & Hausdorf, B. 2020. Changes in the composition of the land snail fauna of Mt. Ciampea, West Java, Indonesia. BIO Web of Conferences, 19: 1–5.

Páll-Gergely, B., Sajan, S., Tripathy, B., Meng, K., Asami, T. & Ablett, J.D. 2020. Genus-level revision of the Alycaeidae (Gastropoda, Cyclophoroidea), with an annotated species catalogue. Zookeys, 981: 1–220.

Purba, C.P.P., Nanggara, S.G., Ratriyono, M., Apriani, I., Rosalina, L., Sari, N.A. & Meridian, A.H. 2014. Potret Keadaan Hutan Indonesia Periode 2009–2013. Bogor: Forest Watch Indonesia [Indonesian].

Régnier, C., Achaz, G., Lambert, A., Cowie, R.H., Bouchet, P. & Fontaine, B. 2015. Mass extinction in poorly known taxa. PNAS, 112(25): 7761–7766.

Reid, W.V. 1998. Biodiversity hotspots. Tree, 13(7): 275–280.

Schilthuizen, M., Liew, T.S., Elahan, B. & Lackman-Ancrenaz, I. 2005. Effect of karst forest degradation on pulmonated and prosobranch land snail communities in Sabah, Malaysian Borneo. Conservation Biology, 19: 949–954.

van Benthem Jutting, W.S.S. 1941. Non-marine mollusca from the satellite islands surrounding Java. Archives Neérlandaises de Zoologie, 5: 251–348.

van Benthem Jutting, W.S.S. 1948. Systematic studies on the non-marine Mollusca of the Indo-Australian archipelago. I. Critical revision of the Javanese pulmonate land-shells of the families Hydrocenidae, Helicinidae, Cyclophoridae, Pupinidae and Cochlostomatidae. Treubia, 19: 539–604.

van Benthem Jutting, W.S.S. 1950. Systematic studies on the non-marine Mollusca of the Indo-Australian archipelago. II. Critical revision of the Javanese pulmonate land-shells of the families Helicarionidae, Pleurodontidae, Fruticicolidae and Streptaxidae. Treubia, 20: 381–505.

van Benthem Jutting, W.S.S. 1952. Systematic studies on the non-marine Mollusca of the Indo-Australian archipelago. III. Critical revision of the Javanese pulmonate land-shells of the families Ellobiidae to Limacidae, with an Appendix on Helicarionidae. Treubia, 21: 291–435.

Vermeulen, J.J. 1996. Notes on terrestrial molluscs of Java, Bali and Nusa Penida. Basteria, 59: 149–162.

Vermeulen, J.J. & Whitten A.J. 1998. Fauna Malesiana. Guide to the Land Snails of Bali. Leiden: Backhuys Publishers.

Whitten, T., Soeriaatmadja, R.E. & Afiff, S.A. 1997. The Ecology of Java and Bali. Hong Kong: Periplus Editions.

Winter, A.J. de. 1983. Euphaedusa cumingiana (L. Pfeiffer) (Pulmonata, Clausiliidae) new to the fauna of Java (Indonesia). Basteria, 47: 59–60.

Winter, A.J. de. & Vermeulen, J.J. 1998. A first record of Truncatellina (Pulmonata: Vertiginidae) from SE. Asia: Truncatellina cf. insulivaga (Pilsbry & Hirase) on Java, Indonesia. Basteria, 62: 269–271.

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